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M.D.
Medical Faculty, Goteborg University, Sweden 1979

Ph.D.
Goteborg University, Sweden 1984

Postdoctoral Fellow
Baylor College of Medicine, 1984-19864

Postdoctoral Fellow
The Swedish Research Council/ Department of Physiology, Göteborg University, Sweden 1987, 1988, 1990

Resident
East Hospital, Göteborg, Sweden 1989-1992